Enumeratio nova Coleopterorum Fennoscandiae, Daniae et Baltiae Hans Silfverberg

Sahlbergia Vol. 9: 1-111, 2004 1 Enumeratio nova Coleopterorum Fennoscandiae, Daniae et Baltiae Hans Silfverberg Silfverberg, H. 2004: Enumeratio nova Coleopterorum Fennoscandiae, Daniae et Baltiae. Sahlbergia 9:1-111. Helsinki, Finland, ISSN 1237-3273. A revised check-list is presented of the Coleoptera of northern Europe (Finland, Sweden, Norway, Denmark, Estonia, Latvia, Lithuania and the Fennoscandian parts of Russia). Hans Silfverberg, Zoological Museum, P.O.Box 17, FI-00014 University of Helsinki; E-mail: hans.silfverberg@helsinki.fi Esipuhe Edellisen pohjoismaisen kovakuoriaisluettelon ilmestymisestä on kulunut toistakymmentä vuotta, ja niin systematiikan kuin faunistiikankin alalla on tapahtunut paljon sen jälkeen. Näin ollen aika on kypsä uutta luetteloa varten. Uuden luettelon järjestys perustuu maailmalla nyt jo laajalti omaksuttuun systeemiin, jonka Lawrence & Newton (1995) esittivät. Uutuudet luettelossa perustuvat pääasiassa jo julkaistuihin tietoihin, Suomen kohdalla on kuitenkin mukana joukko lajeja jotka on ilmoitettu kokouksissa (Liite 1), mutta joiden löydöistä ei vielä ole painettua tietoa. Maat ilmoitetaan samoin kuin ennen, näin ollen F = Suomi, K = Venäjän fennoskandiset osat, S = Ruotsi, N = Norja, D = Tanska, E = Viro, A = Latvia ja I = Liettua (Kuva 1). Erona aikaisempaan vakiintumattomat importtilajit ilmaistaan kyseisen maan pienellä kirjaimella, siten f = importtilaji Suomessa; näissä on pääasiassa huomioitu useita kertoja kulkeutuneet, sekä lähialueilta tulleet lajit. Förord Sedan den föregående nordiska skalbaggskatalogen utkom har över ett decennium gått, och mycket har hänt inom såväl systematik som faunistik. Detta gör att tiden är mogen för en ny katalog. Denna är ordnad enligt det system Lawrence & Newton (1995) uppställt, och som redan accepterats i stora delar av världen. De nya uppgifterna i katalogen baseras i huvudsak på publicerad information, för Finlands del ingår emellertid ett antal arter som anmälts på möten (Appendix 1), men för vilka uppgifter ännu inte kommit i tryck. Länderna anges på samma sätt som förr, d.v.s. F = Finland, K = ryska delarna av Fennoskandien, S = Sverige, N = Norge, D = Danmark, E = Estland, A = Lettland och I = Litauen (Bild 1). En skillnad är att icke etablerade importarter nu anges med liten bokstav för respektive land (t.ex. f = import till Finland); av dessa har huvudsakligen medtagits ofta inkomna arter, samt sådana som kommit från närliggande områden. Introduction A check-list of northern European Coleoptera was published in 1992 (Silfverberg 1992), and after that much has happened both in systematics and in faunistics. A new list is thus appropriate. In general structure this list agrees with the previous one, but some comments on different aspects may be in order. The countries or regions are indicated in the same way as before, that is F = Finland, K = Russian part of Fennoscandia, S = Sweden, N = Norway, D = Denmark, E = Estonia, A = Latvia and I = Lithuania (Figure 1). As a change from the previous arrangement, imported, unestablished species are now indicated with a lower case letter for the area in question (f = imported to Finland, etc.). There has been no attempt to include all imported species; mainly the frequent ones and those imported from nearby areas are mentioned.

2 Silfverberg: Enumeratio Coleopterorum Kuva 1. Luettelossa käytetyt maakohtaiset lyhenteet. Bild 1. De landsvisa förkortningarna som används i katalogen. Figure 1. Abbreviations of the countries as used in the catalogue. The previous list was arranged systematically using the system gradually created by R. Crowson in numerous works. A number of changes were introduced by Lawrence & Newton (1995), and this revised system has already been used widely. It is generally accepted here, too. In some cases the intrafamilial classification is based on still more recent works, so for instance in Adephaga the new Palaearctic catalogue (Löbl & Smetana 2003), in Staphylinidae the catalogue of Herman (2001), in Erotylidae the revision by WÞgrzynowicz (2002) and in Curculionidae to a large extent the catalogue of Alonso-Zarazaga & Lyal (1999). In Staphylinidae many generally used names have been found to be invalid for various reasons (Herman 2001). In order to avoid confusion applications were made (particularly Herman 2002) to the International Commission on Zoological Nomenclature for the preservation a large number of such names, and accordingly customary use has been retained in this catalogue. There are no objective criteria for deciding the extent of genera, and therefore we can see quite different approaches in different works. Recent work in some families has led to heavy splitting, and although such an arrangement can be assumed to give a more detailed picture of systematic relationships, I prefer to have the generic level more equal in different families. Thus the treatment of, for instance, Cerambycidae and Apionidae is relatively conservative. Authors of family-group names were discussed by Lawrence & Newton (1995), and this list generally follows them. Yet many tribal names were not included in their work, and it is quite possible that some of such names will be found to have another author than the one suggested here. The changes to the previous catalogue are primarily based on published papers. Most of them have been mentioned in Silfverberg (1997) or Silfverberg (2003), some further are listed in references below. The status of some Danish species has been changed in accordance with Hansen (1996). There have been many recent advances in Latvian coleopterology, resulting in a check-list of Latvian beetles (Telnov et al. 1997, revised 2002), and the Latvian fauna is mostly listed according to that catalogue. In the previous list there were several species with a B for country, meaning either Estonia or Latvia. Those records that could not be assigned to one country or the other have now been omitted. Concerning the Latvian fauna, the assistance given by A. Barševskis should be specially acknowledged. Some records that have not yet been published are also included. A few species are reported as new to Estonia in a paper by I. Rutanen, soon to be published. A number of species have been reported from Finland in meetings of scientific societies, but have not yet been published in print. They are listed in the Appendix 1, with a reference to the reporter of the species. References Alonso-Zarazaga, M. A. & Lyal, C. H. C. 1999: A world catalogue of families and genera of Curculionoidea (Insecta: Coleoptera) (Except-

Silfverberg: Enumeratio Coleopterorum 3 ing Scolytidae and Platypodidae). Entomopraxis, Barcelona, 315 p. Andersen, J., Hanssen, O. & Ødegaard, F. 2003: Baranowskiella ehnstromi Sörensson, 1997 (Coleoptera, Ptiliidae), the smallest known beetle in Europe, recorded in Norway. Norw. Journ. Entomol. 50:139 141. Assing, V. 2003a: A revision of Calodera Mannerheim. II. A new species, new synonyms, and additional records (Coleoptera: Staphylinidae: Aleocharinae). Beitr. Entomol. 53:217 230. Assing, V. 2003b: On the taxonomy od Gyrohypnus Leach: New synonymies, new species, and a key to the Western palaearctic and Middle Asian representatives of the genus (Insecta: Coleoptera: Staphylinidae). Entomol. Bl. 99:55 81. Barševskis, A. 2002a: Catalogue of Latvian adephagous beetles (Coleoptera: Adephaga). Acta Biol. Univ. Daugavpils 2:1 102. Barševskis, A. 2002b: VaboÝu kârta (Coleoptera). In: Barševskis, A. & al. (eds.) Silenes dabas parka fauna, flora un vegetâcija, Baltijas Koleopterologijas instituts, Daugavpils, p. 37 60. Barševskis, A. 2003: Latvijas skrejvaboýes (Coleoptera: Carabidae, Trachypachidae & Rhysodidae). Baltic Inst. Coleopterol., Daugavpils, 264 p. Barševskis, A., Savenkovs, N. & Valainis, U. 2003: Materiâli par da am retâm un mazpazüstamam vaboýu (Coleoptera) sugâm Ýatvijas faunâ. 2. Internat. Conf. Research and conservation of biological diversity in Baltic Region, Daugavpils. Book of Abstr., p. 19 20. Birkemoe, T. 2002: Lixus iridis (Olivier) (Coleoptera, Curculionidae) in Norway. Norw. Journ. Entomol. 49:153. CibuÝskis, R. 2002a: Distribution and ecology of Noteridae and Dytiscidae (Coleoptera) in the Daugavpils district. Latvijas Entomologs 39:70 78. CibuÝskis, R. 2002b: Distribution and ecology of Oxyteline rove beetles (Coleoptera, Staphylinidae) in Latvia. Latvijas Entomologs 39:80 91. CibuÝskis, R. & Petrova, V. 2002: New species of rove beetles (Coleoptera: Staphylinidae) in fauna of Latvia from Pure. Acta Biol. Univ. Daugavpils 2:103 104. CibuÝskis, R., Barševskis, A. & Leiskina, I. 2003: New species of rove beetles (Coleoptera, Staphylinidae) in Latvian fauna. Baltic Journ. Coleopterol. 3:47 49. Clayhills, T. 2003: Sarvijäärien Stenostola ferrea (Schrank) ja S. dubia (Laicharting) esiintyminen Suomessa (Coleoptera, Cerambycidae). Sahlbergia 7:100. Ericson, B. 2003: Fem för Sverige nya skalbaggar från Malmös närnatur (Coleoptera). Entomol. Tidskr. 124:33 39. Ferrer, J. & Andersson, B. 2002: Förväxling av arterna i släktet Cynaeus Leconte, 1866, i Sverige och Finland [Coleoptera, Tenebrionidae]. Entomol. Tidskr. 123:219 221. Gillerfors, G. 2003a: Monotoma bicolor Villa och Monotoma quadricollis Aubé (Monotomidae) är två skilda arter. Entomol. Tidskr. 124:158. Gillerfors, G. 2003b: Förekomst av knäpparen Negastrius arenicola (Boheman) (Coleoptera, Elateridae) i Sverige. Entomol. Tidskr. 124:173 175. Gønget, H. 2003: The Nemonychidae, Anthribidae and Attelabidae (Coleoptera) of Northern Europe. Fauna Entomol. Scand. 38:1 132. Hansen, M. 1996: Katalog over Danmarks biller. Catalogue of the Coleoptera of Denmark. Entomol. Meddel. 64:1 231. Herman, L. H. 2001: Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Bull. Amer. Mus. Nat. Hist. 265:1 4218. Herman, L. H. 2002: Staphylinidae Latreille, 1804 (Insecta, Coleoptera): proposed conservation of 65 specific names. Bull. Zool. Nomenclat. 59: 99 113. Hyvärinen, E. & Mattila, J. 2004: Carphoborus cholodkowskyi Spessivtseff, 1916, Suomelle uusi kaarnakuoriaislaji (Coleoptera, Scolytidae). Sahlbergia 8:82 83. Jäch, M. A. & Kodada, J. 1999: Parnus obscurus Fabricius is a melandryid! (Coleoptera, Dryopidae, Elmidae, Melandryidae). Entomol. Bl. 95:81 83. Jørum, P., Pedersen, J., Runge, J. B. & Vagtholm- Jensen, O. 2002: Fund af biller i Danmark, 2001. Entomol. Meddel. 70:81 110. Kejval, Z. 2003: The genus Anthelephila (Coleoptera: Anthicidae). Europ. Journ. Entomol. 100:381 392.

4 Silfverberg: Enumeratio Coleopterorum Lawrence, J. F. & Newton, A. F. 1995: Families and subfamilies of Coleoptera (with selected genera, notes, references and data on familygroup names). In: Pakaluk, J. & Slipinski, S.A. (eds.) Biology, Phylogeny and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson, Warszawa, p. 779 1092. Löbl, I. & Smetana, A. (eds.) 2003: Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, 819 p. Miländer, G. 2002a: 28 species of silken fungus beetles (Coleoptera, Cryptophagidae) new to Estonia, from the collection of Valentin Soo. Baltic Journ. Coleopterol. 2:71 75. Miländer, G. 2002b: Anthocomus rufus - uus verevjätklaste (Coleoptera, Malachiidae) liik Eestist. Lepinfo 13:83. Morris, G.M. 2003: An annotated check list of British Curculionoidea (Col.). Entomol. Monthly Mag. 139:193 225. Mossakowski, D., Främbs, H. & Lakomy, W. 2003: The Carabid and Staphilinid fauna of raised bogs. A comparison of Northwest Germany and the Baltic region. Baltic Journ. Coleopterol. 3:137 144. Olberg, S. & Andersen, J. 2003: Contributions to the knowledge of Norwegian Coleoptera. Norw. Journ. Entomol. 50:131 136. Olsvik, P.A., Ødegaard, F. & Hanssen, O. 2001: The beetle fauna (Coleoptera) of the arctic mainland of Norway. Fauna norveg. 21:17 24. Opinion 2053. Staphylinidae Latreille, 1804 (Insecta, Coleoptera): 65 specific names conserved. Bull. Zool. Nomenclat. 60:237 246. Pedersen, J. & Runge, J.B. 2003: Fund af biller i Danmark, 2002 (Coleoptera). Entomol. Meddel. 71:93 113. Rassi, P. 2004: Harpalus rufipalpis Sturm (Carabidae) löydetty Suomesta. Sahlbergia 8:100. Roosileht, U. 2003: New Staphylinidae records for Estonian fauna. Baltic Journ. Coleopterol. 3:35 46. Roosileht, U. & Miländer, G. 2003: Uusi ja haruldasi mardikaliike (Coleoptera) Eesti faunas. Lepinfo (Tartu) 14:100 101. Rutanen, I. 2003: Soukkohärölajien Monotoma quadricollis Aubé ja M. bicolor Villa & Villa esiintymisestä Suomessa (Coleoptera, Monotomidae). Sahlbergia 8:25 26. Silfverberg, H. 1992: Enumeratio Coleopterorum Fennoscandiae, Daniae et Baltiae. Helsingin Hyönteisvaihtoyhdistys - Helsingfors Entomologiska Bytesförening, 94 p. Silfverberg, H. 1997: Additions and corrections to Enumeratio Coleopterorum Fennoscandiae, Daniae et Baltiae. Sahlbergia 3:33 62. Silfverberg, H. 2003: Further additions and corrections to Enumeratio Coleopterorum Fennoscandiae, Daniae et Baltiae. Sahlbergia 7:67 95. Spikkeland, I. & Dolmen, D. 2002: Stenelmis canaliculata (Gyllenhal) (Coleoptera, Elmidae) in Norway. Norw. Journ. Entomol. 49:95 96. Sörensson, M. & Rutanen, I. 2003: Two species of Acrotrichis Motschulsky new to the Finnish fauna (Coleoptera, Ptiliidae) Sahlbergia 7:96 99. Telnov, D. 2002: To the knowledge of Latvian Coleoptera. 2. Latvijas Entomologs 39:16 19. Telnov, D., Barševskis, A., Savich, F., Kovalevsky, F., Berdnikov, S., Doronin, M., Cibulskis, R. & Ratniece, D. 1997: Check-List of Latvian Beetles (Insecta: Coleoptera). Mitt. Internat. Entomol. Ver., Suppl. 5:1 140. Telnov, D., Barševskis, A., Savich, F., Kovalevsky, F., Berdnikov, S., Doronin, M., Cibulskis, R. & Ratniece, D. 2002: The updated list of Latvian beetles (Coleoptera). http://www. lubi.edu.lv/les/coleoptera.htm. Updated March 2003. Telnov, D. & Kalniæš, M. 2003: To the knowledge of Latvian Coleoptera. 3. Latvijas Entomologs 40:21 33. WÞgrzynowicz, P. 2002: Morphology, phylogeny and classification of the family Erotylidae based on adult characters (Coleoptera: Cucujoidea). Genus 13:435 504. Wikars, L.-O., Ahnlund, H. & Wiklund, B. 2004: Atomaria strandi Johnson, en brandgynnad fuktbagge ny för Sverige. - Entomol. Tidskr. 125:57-59. Zamotajlov, A. 2003: Nomenclatural changes in the genus Patrobus (Coleoptera: Carabidae: Patrobini). Acta Soc. Zool. Bohem. 67:239 244.

Silfverberg: Enumeratio Coleopterorum 5 MYXOPHAGA Sphaeriusidae Erichson, 1845 Microsporidae Crotch, 1873 Sphaerius Waltl, 1838 Microsporus Kolenati, 1846 acaroides Waltl, 1838 obsidianus (Kolenati, 1846) ADEPHAGA Gyrinidae Latreille, 1810 Gyrininae Latreille, 1810 Gyrinus Geoffroy, 1762 minutus Fabricius, 1798 opacus Sahlberg, 1819 aeratus Stephens, 1835 pullatus Zaitzev, 1908 marinus Gyllenhal, 1808 colymbus Erichson, 1837 distinctus Aubé, 1836 suffriani Scriba, 1855 natator (Linnaeus, 1758) substriatus Stephens, 1828 urinator Illiger, 1807 caspius Ménétriés, 1832 paykulli Ochs, 1927 Orectochilinae Régimbart, 1882 Orectochilus Dejean, 1833 villosus (Müller, 1776) Haliplidae Brullé, 1835 Brychius Thomson, 1859 elevatus (Panzer, 1793) ssp. elevatus (Panzer, 1793) ssp. norvegicus Munster, 1922 ssp. cristatus J.Sahlberg, 1875 Peltodytes Régimbart, 1879 caesus (Duftschmid, 1805) Haliplus Latreille, 1802 lineatocollis (Marsham, 1802) laminatus (Schaller, 1783) flavicollis Sturm, 1834 fulvus (Fabricius, 1801) lapponum Thomson, 1856 variegatus Sturm, 1834 confinis Stephens, 1828 varius Nicolai, 1822 obliquus (Fabricius, 1787) apicalis Thomson, 1868 FKS-D-AI FKS--EAI F-SNDE-- --SNDEAI --SN---- --S-D--I --S-D--I --S-D--I FK---E-I --SNDE-I furcatus Seidlitz, 1887 F-S-DEAI fulvicollis Erichson, 1837 FKSNDEAfluviatilis Aubé, 1836 lineolatus Mannerheim, 1844 FKSNDEAnomax Balfour-Browne, 1911 interjectus Lindberg, 1937 FK----Asibiricus Motschulsky, 1860 wehnckei Gerhardt, 1877 sahlbergi Falkenström, 1940 lineolatus auct. nec Mannerheim, 1844 ruficollis (DeGeer, 1774) heydeni Wehncke, 1875 immaculatus Gerhardt, 1877 Trachypachidae Thomson, 1857 Trachypachus Motschulsky, 1844 zetterstedtii (Gyllenhal, 1827) FKSN-EA- Noteridae Thomson, 1860 Noterus Clairville, 1806 clavicornis (DeGeer, 1774) crassicornis (Müller, 1776) Dytiscidae Leach, 1815 Copelatinae Van den Branden, 1885 Copelatus Erichson, 1832 haemorrhoidalis (Fabricius, 1787) ruficollis (Schaller, 1783) nec (DeGeer, 1774) Laccophilinae Gistel, 1856 Laccophilus Leach, 1815 hyalinus (DeGeer, 1774) minutus (Linnaeus, 1758) biguttatus Kirby, 1837 strohmi Thomson, 1874 poecilus Klug, 1834 --SN-EAI ponticus Sharp, 1882 variegatus (Germar, 1812) nec (Geoffroy, 1785) obsoletus auct. nec Westhoff, 1881 Hydroporinae Aubé, 1836 Laccornini Wolfe & Roughley, 1990 Laccornis Des Gozis, 1914 oblongus (Stephens, 1835) Hydrovatini Sharp, 1882 Hydrovatus Motschulsky, 1853 cuspidatus (Kunze, 1818) Bidessini Sharp, 1882 Bidessus Sharp, 1882 unistriatus (Goeze, 1777) grossepunctatus Vorbringer, 1907 FKSND-AI

6 Silfverberg: Enumeratio Coleopterorum Hydroglyphus Motschulsky, 1853 Guignotus Houlbert, 1934 geminus (Fabricius, 1792) pusillus (Fabricius, 1781) nec (Müller, 1776) hamulatus (Gyllenhal, 1813) Hyphydrini Sharp, 1882 Hyphydrus Illiger, 1802 ovatus (Linnaeus, 1761) Hygrotini Portevin, 1929 Hygrotus Stephens, 1828 decoratus (Gyllenhal, 1810) inaequalis (Fabricius, 1777) versicolor (Schaller, 1783) quinquelineatus (Zetterstedt, 1828) FKSND-A- Coelambus Thomson, 1860 polonicus (Aubé, 1842) FK---EAI impressopunctatus (Schaller, 1783) parallelogrammus (Ahrens, 1812) F-SNDE-I novemlineatus (Stephens, 1829) marklini (Gyllenhal, 1813) FKSNDEAnigrolineatus (Steven, 1808) F-S-D--- lautus (Schaum, 1843) confluens (Fabricius, 1787) --SNDE-- Hydroporini Aubé, 1836 Hydroporus Clairville, 1806 Hydrocolus Roughley & Nilsson, 2000 nigrita (Fabricius, 1792) brevis F.Sahlberg, 1834 FKSN-EAacutangulus Thomson, 1856 neuter Fairmaire & Laboulbène, 1854 F-SNDEAdiscretus Fairmaire, 1859 pubescens (Gyllenhal, 1808) FKSNDEAfuscipennis Schaum, 1868 planus (Fabricius, 1781) tessellatus (Drapiez, 1819) ------Aobscurus Sturm, 1835 elongatulus Sturm, 1835 eugeniae Zaitzev, 1909 semenowi Jakowlew, 1897 longitarsis J.Sahlberg, 1910 afflatus Scholtz, 1917 tartaricus auct. nec LeConte, 1850 eugeniae auct. nec Zaitzev, 1909 geniculatus Thomson, 1854 subseriatus J.Sahlberg, 1910 pilipes J.Sahlberg, 1921 nigellus Mannerheim, 1853 FKSND-Asubmuticus Thomson, 1874 notabilis LeConte, 1850 ssp. arcticus Thomson, 1856 tomentosus Poppius, 1905 erythrocephalus (Linnaeus, 1758) rufifrons (Müller, 1776) piceus Stephens, 1828 scalesianus Stephens, 1828 angustatus Sturm, 1835 lapponum (Gyllenhal, 1808) puberulus LeConte, 1850 FKS----- levanderi J.Sahlberg, 1889 glabriusculus Aubé, 1838 FKSND-AI umbrosus (Gyllenhal, 1808) morio Aubé, 1838 valliger Hellén, 1929 valliger Hellén, 1929 melanocephalus auct. nec (Marsham, 1802) tristis (Paykull, 1798) gyllenhalii Schiödte, 1841 FKSND--- piceus auct. nec Stephens, 1828 notatus Sturm, 1835 incognitus Sharp, 1869 FKSND-AI striola (Gyllenhal, 1826) palustris (Linnaeus, 1761) neglectus Schaum, 1845 kraatzii Schaum, 1868 -----E-- melanarius Sturm, 1835 longicornis Sharp, 1871 FKSND-AI memnonius Nicolai, 1822 obsoletus Aubé, 1838 --SND--- sahlbergi (Nilsson, 2001) picicornis J.Sahlberg, 1875, nec (Chevrolat, 1863) Porhydrus Guignot, 1945 lineatus (Fabricius, 1775) Graptodytes Seidlitz, 1887 granularis (Linnaeus, 1767) bilineatus (Sturm, 1835) pictus (Fabricius, 1787) Oreodytes Seidlitz, 1887 alpinus (Paykull, 1798) septentrionalis (Gyllenhal, 1826) sanmarkii (Sahlberg, 1826) rivalis (Gyllenhal, 1827) Suphrodytes Des Gozis, 1914 dorsalis (Fabricius, 1787) eljasi (Wirén, 1968) F-S-DEAI FKSND--I

Silfverberg: Enumeratio Coleopterorum 7 Deronectes Sharp, 1882 latus (Stephens, 1829) Scarodytes Des Gozis, 1914 halensis (Fabricius, 1787) Nebrioporus Régimbart, 1906 Potamonectes Zimmermann, 1921 canaliculatus (Lacordaire, 1835) assimilis (Paykull, 1798) depressus (Fabricius, 1775) Stictotarsus Zimmermann, 1919 griseostriatus (DeGeer, 1774) ssp. griseostriatus (DeGeer, 1774) FKSN-E-- ssp. strandi (Brinck, 1943) -K-N---- multilineatus (Falkenström, 1922) duodecimpustulatus (Fabricius, 1792) --SND--- Colymbetinae Erichson, 1837 Agabini Thomson, 1867 Platambus Thomson, 1859 maculatus (Linnaeus, 1758) Agabus Leach, 1817 unguicularis (Thomson, 1867) FKS-DEAaffinis (Paykull, 1798) biguttulus (Thomson, 1867) paludosus (Fabricius, 1801) adpressus Aubé, 1837 didymus (Olivier, 1795) --S-D--I guttatus (Paykull, 1798) melanarius Aubé, 1837 tarsatus Zetterstedt, 1838 bipustulatus (Linnaeus, 1767) solieri Aubé, 1837 conspersus (Marsham, 1802) nebulosus (Forster, 1771) --SNDEAI striolatus (Gyllenhal, 1808) sturmii (Gyllenhal, 1808) arcticus (Paykull, 1798) zetterstedti Thomson, 1856 elongatus (Gyllenhal, 1826) moestus (Curtis, 1835) F------- nigripalpis (J.Sahlberg, 1881) congener (Thunberg, 1794) lapponicus (Thomson, 1867) thomsoni (J.Sahlberg, 1871) pseudoclypealis Scholtz, 1933 FK----Aharaldi Håkan Lindberg, 1933 discolor (Harris, 1828) levanderi Hellén, 1929 approximatus auct. nec Fall, 1923 clypealis (Thomson, 1867) FKS-D-Asetulosus (J.Sahlberg, 1895) FKSN--Aconfinis (Gyllenhal, 1808) FKSN--Ainfuscatus Aubé, 1838 FK-N---- fuscipennis (Paykull, 1798) uliginosus (Linnaeus, 1761) undulatus (Schrank, 1776) --SNDEAI labiatus (Brahm, 1791) serricornis (Paykull, 1799) Ilybius Erichson, 1832 wasastjernai (Sahlberg, 1824) FKSND-Aopacus (Aubé, 1837) chalconatus (Panzer, 1796) F-S-DEAI montanus (Stephens, 1828) ----D-AI melanocornis (Zimmermann, 1915) neclectus (Erichson, 1837) --S-DEAI subtilis (Erichson, 1837) erichsoni (Gemminger & Harold, 1868) nigroaeneus (Erichson, 1837) nec (Marsham, 1802) vittiger (Gyllenhal, 1827) picipes (Kirby, 1837) F-SN---- angustior (Gyllenhal, 1808) crassus Thomson, 1856 FKSND--I subaeneus Erichson, 1837 ater (DeGeer, 1774) guttiger (Gyllenhal, 1808) quadriguttatus (Lacordaire, 1835) obscurus ((Marsham, 1802) similis Thomson, 1856 aenescens Thomson, 1870 fuliginosus (Fabricius, 1792) fenestratus (Fabricius, 1781) Colymbetini Erichson, 1837 Rhantus Dejean, 1833 Nartus Zaitzev, 1907 grapii (Gyllenhal, 1808) suturalis (MacLeay, 1825) pulverosus (Stephens, 1828) frontalis (Marsham, 1802) notatus (Fabricius, 1781) nec (Bergsträsser, 1778) notaticollis (Aubé, 1837) suturellus (Harris, 1828) bistriatus (Bergsträsser, 1778) consputus (Sturm, 1834)

8 Silfverberg: Enumeratio Coleopterorum incognitus Scholz, 1927 ------AI exsoletus (Forster, 1771) latitans Sharp, 1882 -----EAI fennicus Huldén, 1982 FKS----- Colymbetes Clairville, 1806 fuscus (Linnaeus, 1758) paykulli Erichson, 1837 striatus (Linnaeus, 1758) dolabratus (Paykull, 1798) Dytiscinae Leach, 1815 Hydaticini Sharp, 1882 Hydaticus Leach, 1817 transversalis (Pontoppidan, 1763) continentalis Balfour-Browne, 1944 --S-DEAI stagnalis (Fabricius, 1787) nec (Geoffroy, 1785) modestus auct. nec Sharp, 1882 aruspex Clark, 1864 FKSNDEAlaevipennis Thomson, 1867 seminiger (DeGeer, 1774) Aciliini Thomson, 1867 Thermonectini Sharp, 1882 Graphoderus Dejean, 1833 austriacus (Sturm, 1834) --S-D-AI bilineatus (DeGeer, 1774) cinereus (Linnaeus, 1758) zonatus (Hoppe, 1795) ssp. zonatus (Hoppe, 1795) ssp. verrucifer (Sahlberg, 1824) piciventris Thomson, 1868 Acilius Leach, 1817 sulcatus (Linnaeus, 1758) canaliculatus (Nicolai, 1822) Dytiscini Leach, 1815 Dytiscus Linnaeus, 1758 semisulcatus Müller, 1776 dimidiatus Bergsträsser, 1778 marginalis Linnaeus, 1758 latissimus Linnaeus, 1758 circumcinctus Ahrens, 1811 lapponicus Gyllenhal, 1808 circumflexus Fabricius, 1801 Cybistrini Sharp, 1882 Cybister Curtis, 1827 lateralimarginalis (DeGeer, 1774) --SNDEAI --S-DEAI --S-D-AI Carabidae Latreille, 1802 Omophroninae Bonelli, 1810 Omophron Latreille, 1802 limbatum (Fabricius, 1777) --S-DEAI Carabinae Latreille, 1802 Pelophilini Kavanaugh, 1996 Pelophila Dejean, 1821 borealis (Paykull, 1790) FKSN-EA- Nebriini Laporte de Castelnau, 1834 Leistus Frölich, 1799 rufomarginatus (Duftschmid, 1812) --S-D-AI terminatus (Panzer, 1793) rufescens (Fabricius, 1775) nec (Ström, 1768) ferrugineus (Linnaeus, 1758) piceus Frölich, 1799 -----EAI Nebria Latreille, 1802 livida (Linnaeus, 1758) rufescens (Ström, 1768) gyllenhali (Schönherr, 1806) nivalis (Paykull, 1790) brevicollis (Fabricius, 1792) salina Fairmaire & Laboulbène, 1854 --SND--- Notiophilini Motschulsky, 1850 Notiophilus Duméril, 1806 aestuans Dejean, 1826 pusillus Waterhouse, 1833 nec (Schreber, 1759) aquaticus (Linnaeus, 1758) palustris (Duftschmid, 1812) germinyi Fauvel, 1863 stipraisi Barševskis, 1993 rufipes Curtis, 1829 reitteri Spaeth, 1900 biguttatus (Fabricius, 1779) Carabini Latreille, 1802 Calosoma Weber, 1801 sycophanta (Linnaeus, 1758) --S-DE-I inquisitor (Linnaeus, 1758) auropunctatum (Herbst, 1784) --SNDEAI denticolle Gebler, 1833 F------- investigator (Illiger, 1798) reticulatum (Fabricius, 1787) Carabus Linnaeus, 1758 monilis Fabricius, 1792 ---N---- arcensis Herbst, 1784 granulatus Linnaeus, 1758

Silfverberg: Enumeratio Coleopterorum 9 -K---EAI --SND--- FK--DEAI --snd--- -KSND--- --S-DEA- --SNDEAI -----EAI ------A- -K-N---- menetriesi Hummel, 1827 clathratus Linnaeus, 1761 cancellatus Illiger, 1798 ssp. cancellatus Illiger, 1798 rauterbergi Kolbe, 1912 munsteri Born, 1926 ssp. tuberculatus Dejean, 1826 sajanensis Reitter, 1896 kuntzeni Eidam, 1930 auratus Linnaeus, 1761 nemoralis Müller, 1764 hortensis Linnaeus, 1758 glabratus Paykull, 1790 problematicus Herbst, 1786 ssp. harcyniae Sturm, 1815 muelverstedti Reitter, 1896 wockei Born, 1898 relictus Hellén, 1936 ssp. strandi Born, 1926 nitens Linnaeus, 1758 convexus Fabricius, 1775 intricatus Linnaeus, 1761 violaceus Linnaeus, 1758 coriaceus Linnaeus, 1758 Cychrini Laporte de Castelnau, 1834 Cychrus Fabricius, 1794 caraboides (Linnaeus, 1758) Cicindelinae Latreille, 1802 Cicindela Linnaeus, 1758 sylvatica Linnaeus, 1758 hybrida Linnaeus, 1758 maritima Latreille & Dejean, 1822 campestris Linnaeus, 1758 Cylindera Westwood, 1831 germanica (Linnaeus, 1758) arenaria (Fuessly, 1775) ssp. viennensis (Schrank, 1781) Loricerinae Bonelli, 1810 Loricera Latreille, 1802 pilicornis (Fabricius, 1775) Elaphrinae Latreille, 1802 Blethisa Bonelli, 1810 multipunctata (Linnaeus, 1758) Diacheila Motschulsky, 1845 arctica (Gyllenhal, 1810) polita (Faldermann, 1835) Elaphrus Fabricius, 1775 lapponicus Gyllenhal, 1810 uliginosus Fabricius, 1792 cupreus Duftschmid, 1812 riparius (Linnaeus, 1758) aureus Müller, 1821 -----EAI angusticollis F. Sahlberg, 1844 -K----A- Scaritinae Bonelli, 1810 Clivinini Rafinesque, 1815 Clivina Latreille, 1802 fossor (Linnaeus, 1758) collaris (Herbst, 1784) --S-DEAI Dyschirius Bonelli, 1810 digitatus (Dejean, 1825) ------AI thoracicus (Rossi, 1790) arenosus Stephens, 1827 obscurus (Gyllenhal, 1827) angustatus (Ahrens, 1830) nitidus (Dejean, 1825) FK---EAI neresheimeri Wagner, 1915 --S--EAI politus (Dejean, 1825) impunctipennis Dawson, 1854 chalceus Erichson, 1837 --S-DE-- salinus Schaum, 1843 F-SNDEAaeneus (Dejean, 1825) tristis Stephens, 1827 luedersi Wagner, 1915 nigricornis Motschulsky, 1844 FKSN-EAhelleni Müller, 1922 septentrionum Munster, 1923 melancholicus Putzeys, 1866 nigricornis auct. nec Motschulsky, 1844 intermedius Putzeys, 1846 -KS-D-AI laeviusculus Putzeys, 1846 --S-DEAI globosus (Herbst, 1784) Trechinae Bonelli, 1810 Psydrinae LeConte, 1853 Broscini Hope, 1838 Broscus Panzer, 1813 cephalotes (Linnaeus, 1758) Miscodera Eschscholtz, 1830 arctica (Paykull, 1798) Patrobini Kirby, 1837 Patrobus Dejean, 1821 septentrionis Dejean, 1828 australis J.Sahlberg, 1875

10 Silfverberg: Enumeratio Coleopterorum ---N---- --SND-A- ------AI assimilis Chaudoir, 1844 atrorufus (Ström, 1768) Rhysodini Laporte de Castelnau, 1840 Rhysodes Dalman, 1823 sulcatus (Fabricius, 1787) Trechini Bonelli, 1810 Perileptus Schaum, 1860 areolatus (Creutzer, 1799) Aepus Leach, 1819 marinus (Ström, 1788) Trechus Clairville, 1806 secalis (Paykull, 1790) rivularis (Gyllenhal, 1810) rubens (Fabricius, 1792) fulvus Dejean, 1831 quadristriatus (Schranck, 1781) obtusus Erichson, 1837 austriacus Dejean, 1831 Trechoblemus Ganglbauer, 1891 micros (Herbst, 1784) Blemus Dejean, 1821 discus (Fabricius, 1792) Pogonini Laporte de Castelnau, 1834 Pogonus Dejean, 1821 luridipennis (Germar, 1823) chalceus (Marsham, 1802) Bembidiini Stephens, 1827 Bembidiina Stephens, 1827 Asaphidion Des Gozis, 1886 caraboides (Schrank, 1781) pallipes (Duftschmid, 1812) flavipes (Linnaeus, 1761) curtum (Heyden, 1870) austriacum Schweiger, 1975 Bembidion Latreille, 1802 striatum (Fabricius, 1792) velox (Linnaeus, 1761) lapponicum Zetterstedt, 1828 argenteolum Ahrens, 1812 litorale (Olivier, 1791) nigricorne Gyllenhal, 1827 pygmaeum (Fabricius, 1792) lampros (Herbst, 1784) properans (Stephens, 1828) obtusum Audinet-Serville, 1821 -KSN--A- --SN---- --SND--- ----D--I --S-D-A- ------A- FK--DEAI -KSN-EAI FK---EAI --SNDEAI punctulatum Drapiez, 1820 FK---EAI bipunctatum (Linnaeus, 1761) ruficolle (Panzer, 1797) pallidipenne (Illiger, 1802) --SNDEAI illigeri Netolitzky, 1914 F-S-DEAI genei auct. nec Küster, 1847 fellmanni (Mannerheim, 1823) difficile (Motschulsky, 1844) crenulatum F.Sahlberg, 1844 ssp. ponojense J.Sahlberg, 1875 -K------ prasinum (Duftschmid, 1812) hyperboraeorum Munster, 1923 virens Gyllenhal, 1827 hastii Sahlberg, 1827 hirmocaelum Chaudoir, 1850 -K------ tibiale (Duftschmid, 1812) ---N---- mckinleyi Fall, 1926 ssp. scandicum Lindroth, 1943 F-SN---- monticola Sturm, 1825 FK--DEAI deletum Audinet-Serville, 1821 nitidulum (Marsham, 1802) nec (Schrank, 1781) stephensi Crotch, 1869 FKSNDEAlunatum (Duftschmid, 1812) --SND-AI grapii Gyllenhal, 1827 yukonum Fall, 1926 grapeioides Munster, 1930 dauricum (Motschulsky, 1844) F-SN---- tetracolum Say, 1823 ustulatum auct. nec (Linnaeus, 1758) bruxellense Wesmaël, 1835 rupestre auct. nec (Linnaeus, 1758) petrosum Gebler, 1833 ssp. siebkei Sparre Schneider, 1910 ssp. carlhlindrothi Kangas, 1980 -K------ cruciatum Dejean, 1831 andreae auct. nec (Fabricius, 1787) ssp. polonicum J.Müller, 1930 femoratum Sturm, 1825 maritimum (Stephens, 1835) obscurellum (Motschulsky, 1845) -KS-D--- saxatile Gyllenhal, 1827 decorum (Panzer, 1799) dentellum (Thunberg, 1787) tinctum Zetterstedt, 1828 FKS----- varium (Olivier, 1795) semipunctatum (Donovan, 1806) obliquum Sturm, 1825 ephippium (Marsham, 1802)

Silfverberg: Enumeratio Coleopterorum 11 minimum (Fabricius, 1792) normannum Dejean, 1831 azurescens (Dalla Torre, 1877) tenellum Erichson, 1837 articulatum (Panzer, 1796) octomaculatum (Goeze, 1777) doris (Panzer, 1797) schuppelii Dejean, 1831 chaudoirii Chaudoir, 1850 gilvipes Sturm, 1825 fumigatum (Duftschmid, 1812) assimile Gyllenhal, 1810 clarkii (Dawson, 1849) transparens (Gebler, 1829) quadrimaculatum (Linnaeus, 1761) humerale Sturm, 1825 quadripustulatum Audinet-Serville, 1821 biguttatum (Fabricius, 1779) iricolor Bedel, 1879 lunulatum (Geoffroy, 1785) aeneum Germar, 1824 guttula (Fabricius, 1792) neresheimeri Müller, 1929 mannerheimii Sahlberg, 1827 unicolor Chaudoir, 1850 Ocys Stephens, 1828 harpaloides (Audinet-Serville, 1821) quinquestriatus (Gyllenhal, 1810) Cillenus Leach, 1819 lateralis Samouelle, 1819 Tachyina Motschulsky, 1862 Tachys Dejean, 1821 Paratachys Casey, 1918 bistriatus (Duftschmid, 1812) Porotachys Netolitzky, 1914 bisulcatus (Nicolai, 1822) Tachyura Motschulsky, 1862 parvula (Dejean, 1831) Tachyta Kirby, 1837 nana (Gyllenhal, 1810) Harpalinae Bonelli, 1810 Pterostichini Bonelli, 1810 Stomis Clairville, 1806 pumicatus (Panzer, 1796) Poecilus Bonelli, 1810 punctulatus (Schaller, 1783) lepidus (Leske, 1785) kugelanni (Panzer, 1797) dimidiatus (Olivier, 1795) nec (Rossi, 1790) cupreus (Linnaeus, 1758) versicolor (Sturm, 1824) caerulescens auct. nec (Linnaeus, 1758) Pedius Motschulsky, 1850 longicollis (Duftschmid, 1812) inaequalis (Marsham, 1802) nec (Panzer, 1796) Pterostichus Bonelli, 1810 crenatus (Duftschmid, 1812) vernalis (Panzer, 1796) nec (Müller, 1776) macer (Marsham, 1802) ----DEAI aterrimus (Herbst, 1784) aethiops (Panzer, 1796) FK--dEAI madidus (Fabricius, 1775) F---D--- oblongopunctatus (Fabricius, 1787) adstrictus Eschscholtz, 1823 quadrifoveolatus Letzner, 1852 angustatus (Duftschmid, 1812) nec (Fabricius, 1787) niger (Schaller, 1783) melanarius (Illiger, 1798) vulgaris auct. nec (Linnaeus, 1758) nigrita (Paykull, 1790) rhaeticus Heer, 1837 anthracinus (Illiger, 1798) -KS-DEAI gracilis (Dejean, 1828) minor (Gyllenhal, 1827) strenuus (Panzer, 1796) diligens (Sturm, 1824) brevicornis Kirby, 1837 -K------ middendorffi (J.Sahlberg, 1875) -K------ melas (Creutzer, 1799) burmeisteri Heer, 1841 --S-d--- metallicus (Fabricius, 1792) nec (Scopoli, 1763) FKS--EAI --S-DEAI F-S-D-AI -K------ --SNDEAI --S-D--I --S-D-AI FKSND-AI ------AI ---N---- --SNDEA- FK--D-AI FKSND-A- --S-DEAI Abax Bonelli, 1810 parallelepipedus (Piller& Mitterpacher, 1783) --SND--- Sphodrini Laporte de Castelnau, 1834 Calathus Bonelli, 1810 fuscipes (Goeze, 1777) erratus (Sahlberg, 1827) ambiguus (Paykull, 1790) melanocephalus (Linnaeus, 1758) cinctus Motschulsky, 1850 --S-D-Amollis (Marsham, 1802) --SNDEAI ochropterus auct. nec (Duftschmid, 1812) micropterus (Duftschmid, 1812)

12 Silfverberg: Enumeratio Coleopterorum rotundicollis Dejean, 1828 piceus (Marsham, 1802) nec (Linnaeus, 1758) Sphodrus Clairville, 1806 leucophthalmus (Linnaeus, 1758) F-S-DEAI Laemostenus Bonelli, 1810 Pristonychus Dejean, 1828 terricola (Herbst, 1784) Dolichus Bonelli, 1810 halensis (Schaller, 1783) --S-D--I Synuchus Gyllenhal, 1810 vivalis (Illiger, 1798) nivalis (Panzer, 1797) nec (Paykull, 1790) Platyderus Stephens, 1828 depressus (Audinet-Serville, 1821) ---N---- ruficollis (Marsham, 1802) nec (Fabricius, 1801) Platynini Bonelli, 1810 Olisthopus Dejean, 1828 rotundatus (Paykull, 1790) Sericoda Kirby, 1837 bogemannii (Gyllenhal, 1813) FKS----- quadripunctata (DeGeer, 1774) Anchomenus Bonelli, 1810 dorsalis (Pontoppidan, 1763) Oxypselaphus Chaudoir, 1843 obscurus (Herbst, 1784) Paranchus Lindroth, 1974 albipes (Fabricius, 1796) ruficornis (Goeze, 1777) nec (DeGeer, 1774) Platynus Bonelli, 1810 livens (Gyllenhal, 1810) mannerheimii (Dejean, 1828) assimilis (Paykull, 1790) krynickii (Sperk, 1835) --S-DEAI longiventris Mannerheim, 1825 --S--EAI Agonum Bonelli, 1810 scitulum Dejean, 1828 -----EAmicans Nicolai, 1822 piceum (Linnaeus, 1758) gracile Sturm, 1824 latvicum Barševskis, 1993 fuliginosum (Panzer, 1809) thoreyi Dejean, 1828 pelidnum (Paykull, 1790) nec (Herbst, 1784) consimile (Gyllenhal, 1810) munsteri (Hellén, 1935) FKSNDEAexaratum (Mannerheim, 1853) -K------ aldanicum Poppius, 1906 gracilipes (Duftschmid, 1812) marginatum (Linnaeus, 1758) sexpunctatum (Linnaeus, 1758) impressum (Panzer, 1796) -K---EAI ericeti (Panzer, 1809) viridicupreum (Goeze, 1777) -----E-- muelleri (Herbst, 1784) chalconotum Ménétriés, 1832 -K----Asahlbergii (Chaudoir, 1850) archangelicum (J.Sahlberg, 1875) dolens (Sahlberg, 1827) versutum Sturm, 1824 hypocrita (Apfelbeck, 1904) FKS-D-Aviduum (Panzer, 1796) afrum (Duftschmid, 1812) moestum auct. nec (Duftschmid, 1812) duftschmidi Schmidt, 1994 --S-D-AI moestum (Duftschmid, 1812) nec (Gmelin, 1790) lugens (Duftschmid, 1812) F-S-DEAI Zabrini Bonelli, 1810 Amarina Zimmermann, 1831 Amara Bonelli, 1810 strenua Zimmermann, 1832 ----D-Atricuspidata Dejean, 1831 plebeja (Gyllenhal, 1810) concinna Zimmermann, 1832 similata (Gyllenhal, 1810) ovata (Fabricius, 1792) montivaga Sturm, 1825 nitida Sturm, 1825 communis (Panzer, 1797) pulpani Kult, 1949 ------Apseudocommunis Burakowski, 1957 convexior Stephens, 1828 FKSND-AI nigricornis Thomson, 1857 lunicollis Schiödte, 1837 curta Dejean, 1828 littorea Thomson, 1857 aenea (DeGeer, 1774) spreta Dejean, 1831 famelica Zimmermann, 1832 eurynota (Panzer, 1796) familiaris (Duftschmid, 1812) anthobia Villa & Villa, 1833 --S-D-Alucida (Duftschmid, 1812)

Silfverberg: Enumeratio Coleopterorum 13 tibialis (Paykull, 1798) erratica (Duftschmid, 1812) interstitialis Dejean, 1828 ingenua (Duftschmid, 1812) fusca Dejean, 1828 cursitans Zimmermann, 1832 municipalis (Duftschmid, 1812) bifrons (Gyllenhal, 1810) infima (Duftschmid, 1812) praetermissa (Sahlberg, 1827) brunnea (Gyllenhal, 1810) quenseli (Schönherr, 1806) crenata Dejean, 1828 F-S-D-AI apricaria (Paykull, 1790) fulva (Müller, 1776) majuscula (Chaudoir, 1850) consularis (Duftschmid, 1812) aulica (Panzer, 1796) gebleri Dejean, 1831 helleri Gredler, 1868 convexiuscula (Marsham, 1802) FKSNDEAtorrida (Panzer, 1796) alpina (Paykull, 1790) hyperborea Dejean, 1831 equestris (Duftschmid, 1812) Zabrina Bonelli, 1810 Zabrus Clairville, 1806 tenebrioides (Goeze, 1777) Panagaeini Bonelli, 1810 Panagaeus Latreille, 1802 cruxmajor (Linnaeus, 1758) bipustulatus (Fabricius, 1775) Chlaeniini Brullé, 1834 Chlaenius Bonelli, 1810 tristis (Schaller, 1783) spoliatus (Rossi, 1790) nigricornis (Fabricius, 1787) nitidulus (Schrank, 1781) tibialis Dejean, 1826 ------Akindermanni auct. nec Chaudoir, 1856 vestitus (Paykull, 1790) --S-D-AI sulcicollis (Paykull, 1798) F-S-DEAI quadrisulcatus (Paykull, 1790) -KS-DEAcostulatus (Motschulsky, 1859) FKS--EAI Oodini LaFerté-Sénectère, 1851 Oodes Bonelli, 1810 helopioides (Fabricius, 1792) gracilis Villa & Villa, 1833 --S--EAI Licinini Bonelli, 1810 Badister Clairville, 1806 unipustulatus Bonelli, 1813 bullatus (Schrank, 1798) bipustulatus (Fabricius, 1792) nec (Fabricius, 1775) meridionalis Puel, 1925 --S-DEAI lacertosus Sturm, 1815 sodalis (Duftschmid, 1812) -KSNDEAI dorsiger (Duftschmid, 1812) ----D-Apeltatus (Panzer, 1796) dilatatus Chaudoir, 1837 collaris Motschulsky, 1844 --S-DEAI anomalus (Perris, 1866) striatulus Hansen, 1944 Licinus Latreille, 1802 depressus (Paykull, 1790) Harpalini Bonelli, 1810 Harpalina Bonelli, 1810 Ophonus Stephens, 1828 rupicola (Sturm, 1818) --S-D-Alaticollis (Mannerheim, 1825) nitidulus Stephens, 1828 punctatulus (Duftschmid, 1812) nec (Fabricius, 1792) puncticollis (Paykull, 1798) melletii (Heer, 1837) rufibarbis (Fabricius, 1792) seladon (Schauberger, 1926) schaubergerianus (Puel, 1937) F-----Apuncticeps Stephens, 1828 -KS-D-Aazureus (Fabricius, 1775) --S-DEA- Harpalus Latreille, 1802 Pseudoophonus Motschulsky, 1844 Semiophonus Schauberger, 1933 griseus (Panzer, 1796) rufipes (DeGeer, 1774) pubescens (Müller, 1776) calceatus (Duftschmid, 1812) melancholicus Dejean, 1829 signaticornis (Duftschmid, 1812) --S-DE-I affinis (Schrank, 1781) aeneus (Fabricius, 1775) nec (DeGeer, 1774) distinguendus (Duftschmid, 1812) smaragdinus (Duftschmid, 1812) caspius (Steven, 1806) roubali Schauberger, 1928 serripes (Quensel, 1806) --S-D--I

14 Silfverberg: Enumeratio Coleopterorum autumnalis (Duftschmid, 1812) --S---AI atratus Latreille, 1804 solitaris Dejean, 1829 fuliginosus (Duftschmid, 1812) nec (Panzer, 1809) latus (Linnaeus, 1758) nigritarsis Sahlberg, 1827 FKS----- luteicornis (Duftschmid, 1812) xanthopus Gemminger & Harold, 1868 ssp. winkleri Schauberger, 1923 laevipes Zetterstedt, 1828 quadripunctatus Dejean, 1829 rubripes (Duftschmid, 1812) progrediens Schauberger, 1922 -----EArufipalpis Sturm, 1818 F-S-DEAI rufitarsis (Duftschmid, 1812) nec (Illiger, 1802) neglectus Audinet-Serville, 1821 --S-D--I servus (Duftschmid, 1812) --S-D-AI tardus (Panzer, 1796) modestus Dejean, 1829 anxius (Duftschmid, 1812) picipennis (Duftschmid, 1812) --S-DEAI pumilus Sturm, 1818 --S-D-AI vernalis (Fabricius, 1801) nec (Panzer, 1796) froelichii Sturm, 1818 -KS-DEAI hirtipes (Panzer, 1797) --S-DEAI flavescens (Piller & Mitterpacher, 1783) FKS-D-AI rufus Brüggemann, 1873 Microderes Faldermann, 1835 brachypus (Steven, 1809) Anisodactylina Lacordaire, 1854 Anisodactylus Dejean, 1829 poeciloides (Stephens, 1828) binotatus (Fabricius, 1787) nemorivagus (Duftschmid, 1812) signatus (Panzer, 1797) Diachromus Erichson, 1837 germanus (Linnaeus, 1758) Stenolophina Kirby, 1837 Stenolophus Dejean, 1821 teutonus (Schrank, 1781) skrimshiranus Stephens, 1828 mixtus (Herbst, 1784) Bradycellus Erichson, 1837 Tetraplatypus Tschitschérine, 1897 ruficollis (Stephens, 1828) similis (Dejean, 1829) verbasci (Duftschmid, 1812) F------- FK--D--I ----D-AI ----D-AI --S-D-AI F-SND--- harpalinus (Audinet-Serville, 1821) -KSNDEAI csikii Laczó, 1912 F-S-D--- caucasicus (Chaudoir, 1846) collaris (Paykull, 1798) nec (Herbst, 1784) Dicheirotrichus Jacquelin du Val, 1857 Trichocellus Ganglbauer, 1892 gustavii Crotch, 1871 -KSND--- pubescens (Paykull, 1790) nec (Müller, 1776) mannerheimii (F.Sahlberg, 1844) -K------ rufithorax (Sahlberg, 1827) cognatus (Gyllenhal, 1827) placidus (Gyllenhal, 1827) Acupalpus Dejean, 1829 flavicollis (Sturm, 1825) brunnipes (Sturm, 1825) --SN-E-I meridianus (Linnaeus, 1761) suturalis Dejean, 1829 ------Aparvulus (Sturm, 1825) dorsalis (Fabricius, 1787) nec (Pontoppidan, 1763) dubius Schilsky, 1888 --S-DEAexiguus Dejean, 1829 elegans (Dejean, 1829) interstitialis Reitter, 1884 -----EA- Anthracus Motschulsky, 1850 consputus (Duftschmid, 1812) F-S-DEAI Perigonini Horn, 1881 Perigona Laporte de Castelnau, 1835 Trechicus LeConte, 1853 nigriceps (Dejean, 1831) F-SND-A- Cyclosomini Laporte de Castelnau, 1834 Masoreus Dejean, 1821 wetterhallii (Gyllenhal, 1813) --SNDEAI Odacanthini Laporte de Castelnau, 1834 Odacantha Paykull, 1798 melanura (Linnaeus, 1767) Lebiini Bonelli, 1810 Lebiina Bonelli, 1810 Lebia Latreille, 1802 cyanocephala (Linnaeus, 1758) chlorocephala (Hoffmannsegg, 1803) cruxminor (Linnaeus, 1758) Demetriina Bates, 1886 Demetrias Bonelli, 1810 monostigma Samouelle, 1819 F-S-DEAI atricapillus (Linnaeus, 1758) --S-D-Aimperialis (Germar, 1824)

Silfverberg: Enumeratio Coleopterorum 15 Dromiina Bonelli, 1810 Paradromius Fowler, 1887 longiceps (Dejean, 1826) F-S-DEAlinearis (Olivier, 1795) Dromius Bonelli, 1810 agilis (Fabricius, 1787) angustus Brullé, 1834 --SND--- quadraticollis Morawitz, 1862 FKS--EAI meridionalis Dejean, 1825 schneideri Crotch, 1871 marginellus (Fabricius, 1794) nec (Herbst, 1784) fenestratus (Fabricius, 1794) quadrimaculatus (Linnaeus, 1758) Calodromius Reitter, 1905 spilotus (Illiger, 1798) quadrinotatus (Panzer, 1800) nec (Fabricius, 1798) F-SNDEA- Philorhizus Hope, 1838 quadrisignatus (Dejean, 1825) sigma (Rossi, 1790) melanocephalus (Dejean, 1825) notatus (Stephens, 1827) nigriventris (Thomson, 1857) Microlestes Schmidt-Goebel, 1846 minutulus (Goeze, 1777) maurus (Sturm, 1827) Lionychina Jeannel, 1948 Syntomus Hope, 1838 truncatellus (Linnaeus, 1761) ai Barševskis, 1993 foveatus (Geoffroy, 1785) Lionychus Wissmann, 1846 quadrillum (Duftschmid, 1812) F-S----- Cymindina Laporte de Castelnau, 1834 Cymindis Latreille, 1806 humeralis (Geoffroy, 1785) --S-D-AI angularis Gyllenhal, 1810 macularis Mannerheim, 1824 vaporariorum (Linnaeus, 1758) Brachininae Bonelli, 1810 Brachinus Weber, 1801 crepitans (Linnaeus, 1758) F-S-DEA- POLYPHAGA HYDROPHILOIDEA Hydrophilidae Latreille, 1802 Helophorinae Leach, 1815 Helophorus Fabricius, 1775 nubilus Fabricius, 1777 FK--DEAI F-SND-A- FK--DEAI FK---E-- F-SND--- --S-DE-I FKSN-E-- --SND--- tuberculatus Gyllenhal, 1808 sibiricus (Motschulsky, 1860) fennicus auct. nec Paykull, 1798 aquaticus (Linnaeus, 1758) aequalis Thomson, 1868 grandis Illiger, 1798 strandi Angus, 1970 brevipalpis Bedel, 1881 arvernicus Mulsant, 1846 glacialis Villa, 1833 granularis (Linnaeus, 1761) discrepans Rey, 1885 minutus Fabricius, 1775 lapponicus Thomson, 1853 fulgidicollis Motschulsky, 1860 griseus Herbst, 1793 nanus Sturm, 1836 redtenbacheri Kuwert, 1885 pumilio Erichson, 1837 pallidus Gebler, 1830 laticollis Thomson, 1853 strigifrons Thomson, 1868 asperatus Rey, 1885 flavipes Fabricius, 1792 obscurus Mulsant, 1844 walkeri Sharp, 1916 Georissinae Laporte de Castelnau, 1840 Georissus Latreille, 1809 crenulatus (Rossi, 1794) Hydrochinae Thomson, 1859 Hydrochus Leach, 1817 elongatus (Schaller, 1783) F-S-D-Aignicollis Motschulsky, 1860 elongatus auct. nec (Schaller, 1783) crenatus (Fabricius, 1792) F-S-DEAI carinatus Germar, 1824 brevis (Herbst, 1793) megaphallus van Berge Henegouwen, 1988 F-SNDE-- angustatus Germar, 1824 -----EA- Spercheinae Erichson, 1837 Spercheus Illiger, 1798 emarginatus (Schaller, 1783) -KS-DEAI Hydrophilinae Latreille, 1802 Berosini Mulsant, 1844 Berosus Leach, 1817 spinosus (Steven, 1808) fulvus Kuwert, 1888 F-SNDE--

16 Silfverberg: Enumeratio Coleopterorum signaticollis (Charpentier, 1825) luridus (Linnaeus, 1761) Chaetarthriini Bedel, 1881 (1844) Chaetarthria Stephens, 1832 seminulum (Herbst, 1797) Anacaenini Hansen, 1991 Paracymus Thomson, 1867 aeneus (Germar, 1824) Anacaena Thomson, 1859 globulus (Paykull, 1798) lutescens (Stephens, 1829) limbata auct. nec (Fabricius, 1792) limbata (Fabricius, 1792) Laccobiini Bertrand, 1954 Oocyclini Hansen, 1991 Laccobius Erichson, 1837 decorus (Gyllenhal, 1827) minutus (Linnaeus, 1758) colon (Stephens, 1829) biguttatus Gerhardt, 1877 sinuatus Motschulsky, 1849 striatulus (Fabricius, 1801) bipunctatus (Fabricius, 1775) Hydrophilini Latreille, 1802 Helochares Mulsant, 1844 punctatus Sharp, 1869 lividus auct. nec (Forster, 1771) obscurus (Müller, 1776) griseus auct. nec (Fabricius, 1787 Enochrus Thomson, 1859 melanocephalus (Olivier, 1792) ochropterus (Marsham, 1802) frontalis (Erichson, 1837) fuscipennis (Thomson, 1884) sahlbergi (Fauvel, 1887) quadripunctatus (Herbst, 1797) halophilus (Bedel, 1878) bicolor (Fabricius, 1792) testaceus (Fabricius, 1801) affinis (Thunberg, 1794) coarctatus (Gredler, 1863) Cymbiodyta Bedel, 1881 marginella (Fabricius, 1792) Hydrobius Leach, 1815 fuscipes (Linnaeus, 1758) subrotundus Stephens, 1829 rottenbergi Gerhardt, 1872 F-S?DE-I --SND--- F-SNDEA- F-S-D-A- F-S--E-- FKSND-A- FKS-D--I -KSNDEAI ----D--I FKSND-AI arcticus Kuwert, 1890 Limnoxenus Motschulsky, 1853 niger (Gmelin, 1790) Hydrochara Berthold, 1827 caraboides (Linnaeus, 1758) flavipes (Steven, 1808) Hydrophilus Geoffroy, 1762 Hydrous Linnaeus, 1775 piceus (Linnaeus, 1758) aterrimus Eschscholtz, 1822 F-SND-AI ------A- Sphaeridiinae Latreille, 1802 Coelostomatini Heyden, Reitter & Weise, 1891 Coelostoma Brullé, 1835 orbiculare (Fabricius, 1775) Megasternini Mulsant, 1844 Cercyon Leach, 1817 ustulatus (Preyssler, 1790) laminatus Sharp, 1873 F-S-DEAI littoralis (Gyllenhal, 1808) depressus Stephens, 1829 obsoletus (Gyllenhal, 1808) lugubris auct. nec (Olivier, 1790) impressus (Sturm, 1807) haemorrhoidalis (Fabricius, 1775) melanocephalus (Linnaeus, 1758) emarginatus Baranowski, 1985 F-SN---- borealis Baranowski, 1985 F-SN---- lateralis (Marsham, 1802) bifenestratus Küster, 1851 marinus Thomson, 1853 unipunctatus (Linnaeus, 1758) quisquilius (Linnaeus, 1761) terminatus (Marsham, 1802) pygmaeus (Illiger, 1801) nigriceps (Marsham, 1802) FKSNDEAatricapillus (Marsham, 1802) granarius Erichson, 1837 -K--D-AI tristis (Illiger, 1801) convexiusculus Stephens, 1829 sternalis Sharp, 1918 F-S-D-Aanalis (Paykull, 1798) Megasternum Mulsant, 1844 concinnum (Marsham, 1802) obscurum (Marsham, 1802) Cryptopleurum Mulsant, 1844 subtile Sharp, 1884 F-SND---

Silfverberg: Enumeratio Coleopterorum 17 minutum (Fabricius, 1775) crenatum (Panzer, 1794) Sphaeridiini Latreille, 1802 Sphaeridium Fabricius, 1775 bipustulatum Fabricius, 1781 marginatum Fabricius, 1787 F-SND--I substriatum Faldermann, 1838 --S-D--I lunatum Fabricius, 1792 scarabaeoides (Linnaeus, 1758) HISTEROIDEA Sphaeritidae Shuckard, 1839 Sphaerites Duftschmid, 1805 glabratus (Fabricius, 1792) Histeridae Gyllenhal, 1808 Abraeinae MacLeay, 1819 Abraeini MacLeay, 1819 Chaetabraeus Portevin, 1929 globulus (Creutzer, 1799) --S-DE-I Abraeus Leach, 1817 granulum Erichson, 1839 perpusillus (Marsham, 1802) --S-DEAI globosus (Hoffmann, 1803) parvulus Aubé, 1842 Plegaderini Portevin, 1929 Plegaderus Erichson, 1834 saucius Erichson, 1834 vulneratus (Panzer, 1797) caesus (Herbst, 1792) F-SN--AI dissectus Erichson, 1839 --S-D--I sanatus Truqui, 1852 ssp. gobanzi Müller, 1902 Acritini Wenzel, 1944 Acritus LeConte, 1853 minutus (Herbst, 1792) nigricornis (Hoffmann, 1803) FKSNDEAkomai Lewis, 1879 homoeopathicus Wollaston, 1857 ---ND-A- Aeletes Horn, 1873 atomarius (Aubé, 1842) Teretriini Bickhardt, 1914 Teretrius Erichson, 1834 fabricii Mazur, 1972 F-S--EAI picipes (Fabricius, 1792) nec (Olivier, 1789) Saprininae Blanchard, 1845 Saprinus Erichson, 1834 rugifer (Paykull, 1809) FKSND-AI quadristriatus auct. nec (Thunberg, 1794) ----D-A- ------A- --S-DEAI planiusculus Motschulsky, 1849 cuspidatus Ihssen, 1949 semistriatus (Scriba, 1790) subnitescens Bickhardt, 1909 tenuistrius Marseul, 1855 virescens (Paykull, 1798) immundus (Gyllenhal, 1827) aeneus (Fabricius, 1775) Chalcionellus Reichardt, 1932 decemstriatus (Rossi, 1792) conjungens (Paykull, 1798) Hypocacculus Bickhardt, 1916 rufipes (Kugelann, 1792) Hypocaccus Thomson, 1867 Baeckmanniolus Reichardt, 1926 rugiceps (Duftschmid, 1805) metallicus (Herbst, 1792) rugifrons (Paykull, 1798) dimidiatus (Illiger, 1807) ssp. maritimus (Stephens, 1830) Gnathoncus Jacquelin du Val, 1858 rotundatus (Kugelann, 1792) --S-DEAI --S-D-AI FKSND-AI --SNDEAI nanus (Scriba, 1790) nec (Piller& Mitterpacher, 1783) punctulatus Thomson, 1862 nannetensis (Marseul, 1862) communis (Marseul, 1862) FKSND--- schmidti Reitter, 1894 buyssoni Auzat, 1917 FKSND--I nidorum Stockmann, 1957 F-S---AI Myrmetes Marseul, 1862 paykulli Kanaar, 1979 piceus (Paykull, 1809) nec (Marsham, 1802) Dendrophilinae Reitter, 1909 Dendrophilini Reitter, 1909 Dendrophilus Leach, 1817 corticalis (Paykull, 1798) punctatus (Herbst, 1792) nec (Müller, 1776) pygmaeus (Linnaeus, 1758) Paromalini Reitter, 1909 Carcinops Marseul, 1855 pumilio (Erichson, 1834) quatuordecimstriata (Stephens, 1835) Platylomalus Cooman, 1948 complanatus (Panzer, 1797) -K-N--A-

18 Silfverberg: Enumeratio Coleopterorum F-SNDEA- FKSNdEAI --S-DE-- F-S-DEAI ----D-AI -KSNDEAI --S-DEA- -----EA- ----DEAI -----E-- FK---EAI ----DE-- Paromalus Erichson, 1834 Microlomalus Lewis, 1907 flavicornis (Herbst, 1792) parallelepipedus (Herbst, 1792) Onthophilinae MacLeay, 1819 Onthophilus Leach, 1817 striatus (Forster, 1771) punctatus (Müller, 1776) sulcatus (Moll, 1784) Histerinae Gyllenhal, 1808 Histerini Gyllenhal, 1808 Margarinotus Marseul, 1853 Paralister Bickhardt, 1917 Stenister Reichardt, 1926 Grammostethus Lewis, 1906 brunneus (Fabricius, 1775) impressus (Fabricius, 1798) cadaverinus (Hoffmann, 1803) striola (Sahlberg, 1819) succicola (Thomson, 1862) merdarius (Hoffmann, 1803) terricola (Germar, 1824) bipustulatus (Schrank, 1781) obscurus (Kugelann, 1792) stercorarius (Hoffmann, 1803) ventralis (Marseul, 1854) neglectus (Germar, 1813) purpurascens (Herbst, 1792) carbonarius (Hoffmann, 1803) marginatus (Erichson, 1834) Hister Linnaeus, 1758 quadrimaculatus Linnaeus, 1758 unicolor Linnaeus, 1758 quadrinotatus Scriba, 1790 illigeri Duftschmid, 1805 uncinatus Illiger, 1807 helluo Truqui, 1852 bissexstriatus Fabricius, 1801 funestus Erichson, 1834 Atholus Thomson, 1859 bimaculatus (Linnaeus, 1758) duodecimstriatus (Schrank, 1781) praetermissus (Peyron, 1856) corvinus (Germar, 1817) Platysomatini Bickhardt, 1914 Platysoma Leach, 1817 Eurylister Bickhardt, 1920 Cylister Cooman, 1941 minus (Rossi, 1792) frontale (Paykull, 1798) compressum (Herbst, 1783) FKS-D-AI deplanatum (Gyllenhal, 1808) angustatum (Hoffmann, 1803) ferrugineum (Thunberg, 1794) nec (Olivier, 1789) lineare Erichson, 1834 elongatum (Thunberg, 1787) FKS---AI oblongum (Fabricius, 1792) ------A- Hololeptini Hope, 1840 Hololepta Paykull, 1811 plana (Sulzer, 1776) Hetaeriinae Marseul, 1857 Hetaerius Erichson, 1834 ferrugineus (Olivier, 1789) STAPHYLINOIDEA Hydraenidae Mulsant, 1844 Hydraeninae Mulsant, 1844 Limnebius Leach, 1815 truncatellus (Thunberg, 1794) papposus Mulsant, 1844 crinifer Rey, 1885 parvulus (Herbst, 1797) truncatulus Thomson, 1853 nitidus (Marsham, 1802) picinus (Marsham, 1802) aluta Bedel, 1881 Hydraena Kugelann, 1794 testacea Curtis, 1830 palustris Erichson, 1837 britteni Joy, 1907 riparia Kugelann, 1794 reyi Kuwert, 1888 sternalis auct. nec Rey, 1893 nigrita Germar, 1824 pulchella Germar, 1824 gracilis Germar, 1824 excisa Kiesenwetter, 1849 Ochthebiinae Thomson, 1859 Ochthebius Leach, 1815 auriculatus Rey, 1886 dilatatus Stephens, 1829 FK-NDEAI --S-D-AI --S-D-AI FKS-DE-I F-S-DE-I FKSND--- --SNDE-- FKS-DEA- F-----A-

Silfverberg: Enumeratio Coleopterorum 19 bicolon Germar, 1824 F-SND--- czwalinai Kuwert, 1887 F------- stockmanni Balfour-Browne, 1948 kaninensis Poppius, 1909 F------- minimus (Fabricius, 1792) impressus (Marsham, 1802) remotus Reitter, 1887 ------Aalpinus (Ieniåtea, 1979) rugulosus auct. nec (Wollaston, 1857) hungaricus Endrödy-Younga, 1967 FK---E-- nilssoni Hebauer, 1986 metallescens Rosenhauer, 1847 pusillus Stephens, 1835 marinus (Paykull, 1798) FKSNDEAlenensis Poppius, 1907 -K-N---- viridis Peyron, 1858 evanescens J.Sahlberg, 1875 -K------ Ptiliidae Heer, 1843 Ptiliinae Heer, 1843 Nossidium Erichson, 1845 pilosellum (Marsham, 1802) Ptenidium Erichson, 1845 gressneri Erichson, 1845 --S-D-AI laevigatum Erichson, 1845 F-SND-Aturgidum Thomson, 1855 --SND-AI intermedium Wankowicz, 1869 --SND-AI fuscicorne Erichson, 1845 FKSND--I punctatum (Gyllenhal, 1827) F-SNDE-- formicetorum Kraatz, 1851 FKSNDEAmyrmecophilum (Motschulsky, 1845) nec (Allibert, 1844) pusillum (Gyllenhal, 1808) longicorne (Fuss, 1868) ------Anitidum (Heer, 1841) Actidium Matthews, 1868 boudieri (Allibert, 1844) ----D-Acoarctatum (Haliday, 1855) Oligella Motschulsky, 1868 foveolata (Allibert, 1844) FKSND-Anana (Strand, 1946) --SN---- intermedia Besuchet, 1971 Micridium Motschulsky, 1868 angulicolle (Fairmaire, 1857) halidaii (Matthews, 1868) F-SND-A- Ptilium Gyllenhal, 1827 Millidium Motschulsky, 1855 minutissimum (Ljungh, 1804) affine Erichson, 1845 caesum Erichson, 1845 myrmecophilum (Allibert, 1844) exaratum (Allibert, 1844) horioni Rosskothen, 1934 modestum Wankowicz, 1869 Euryptilium Matthews, 1872 gillmeisteri Flach, 1889 saxonicum (Gillmeister, 1845) marginatum auct. nec (Aubé, 1850) Ptiliola Haldeman, 1848 Nanoptilium Flach, 1889 kunzei (Heer, 1841) brevicollis (Matthews, 1860) Ptiliolum Flach, 1888 stockmanni Besuchet, 1971 caledonicum (Sharp, 1871) croaticum (Matthews, 1872) schwarzi (Flach, 1887) wuesthoffi Rosskothen, 1934 spencei (Allibert, 1844) fuscum (Erichson, 1845) flachi (Reitter, 1909) marginatum (Aubé, 1850) lederi (Flach, 1888) sahlbergi (Flach, 1888) Microptilium Matthews, 1872 palustre Kuntzen, 1914 Nanosellinae Barber, 1924 Baranowskiella Sörensson, 1997 ehnstromi Sörensson, 1997 Acrotrichinae Reitter, 1909 (1856) Ptinella Motschulsky, 1844 limbata (Heer, 1841) testacea (Heer, 1841) denticollis (Fairmaire, 1858) johnsoni Rutanen, 1985 aptera (Guérin-Ménéville, 1839) britannica Matthews, 1858 tenella (Erichson, 1845) microscopica (Gillmeister, 1845) Pteryx Matthews, 1858 suturalis (Heer, 1841) splendens Strand, 1960 Nephanes Thomson, 1859 titan (Newman, 1834) -KS-D--- FKSND-A- -KSND--- F-SND--I ---ND-A- FKSND--- F-SND-A- F------- FKSN-E-- F-SNDE-- F-SND--- F-SNDE-- F-SND--- FKSN-EA- F-SN---- F-SND--I FKS-D--- F-S----- F-SND--- FKSND--- FKS----- FKSND-A- F-SN---- FKSND-AI

20 Silfverberg: Enumeratio Coleopterorum Smicrus Matthews, 1872 filicornis (Fairmaire& Laboulbène, 1855) FKSND-A- Baeocrara Thomson, 1859 variolosa (Mulsant & Rey, 1873) FKSND--I littoralis (Thomson, 1855) nec (Motschulsky, 1845) japonica (Matthews, 1884) F-SND-A- Acrotrichis Motschulsky, 1848 grandicollis (Mannerheim, 1844) montandonii (Allibert, 1844) chevrolatii (Allibert, 1844) FKSND--- thoracica (Waltl, 1838) FKSND-Asericans (Heer, 1841) FKSNDEApicicornis auct. nec (Mannerheim, 1843) dispar (Matthews, 1865) brevipennis (Erichson, 1845) FKSNDEApumila (Erichson, 1845) longicornis auct. nec (Mannerheim, 1844) silvatica Rosskothen, 1935 FKSND--- parva Rosskothen, 1935 FKSND-Avolans (Motschulsky, 1845) fennica Renkonen, 1939 cognata (Matthews, 1877) platonoffi Renkonen, 1945 insularis (Mäklin, 1852) F-SND--- norvegica Strand, 1941 FKSND--- arnoldi Rosskothen, 1935 intermedia (Gillmeister, 1845) rosskotheni Sundt, 1971 F-SND--- fraterna Johnson, 1975 henrici (Matthews, 1872) ---ND--- fratercula (Matthews, 1878) atomaria (DeGeer, 1774) lucidula Rosskothen, 1935 F-SND--- danica Sundt, 1958 sitkaensis (Motschulsky, 1845) FKSNDEAfratercula auct. nec (Matthews, 1878) sjobergi Sundt, 1958 F-SN---- fascicularis (Herbst, 1793) rugulosa Rosskothen, 1935 strandi Sundt, 1958 FKSND-Asuecica Sundt, 1958 --SND--- Actinopteryx Matthews, 1872 fucicola (Allibert, 1844) Agyrtidae Thomson, 1859 Pterolominae Thomson, 1862 Pteroloma Gyllenhal, 1827 forsstromii (Gyllenhal, 1810) FKSN-E-- Agyrtinae Thomson, 1859 Ecanus Stephens, 1833 glaber (Fabricius, 1787) Agyrtes Frölich, 1799 bicolor Laporte de Castelnau, 1840 Leiodidae Fleming, 1821 Leiodinae Fleming, 1821 Sogdini Lopatin, 1961 Triarthron Märkel, 1840 maerkelii Märkel, 1840 Sogda Lopatin, 1961 Trichohydnobius Vogt, 1961 suturalis (Zetterstedt, 1828) perrisii (Fairmaire, 1855) ciliaris (Thomson, 1874) hyperborea (Strand, 1943) F-S----- FKSND--- FKSND--I FK--D-A- F--N---- FKSN-E-- F------- --SN---- Hydnobius Schmidt, 1841 septentrionalis Thomson, 1874 punctatus (Sturm, 1807) latifrons (Curtis, 1840) strigosus Schmidt, 1841 tibialis J.Sahlberg, 1903 spinipes (Gyllenhal, 1813) edentatus (Sahlberg, 1833) orientalis Hlisnikovsky, 1967 claviger Strand, 1943 multistriatus (Gyllenhal, 1813) Leiodini Fleming, 1821 Leiodes Latreille, 1796 punctulata (Gyllenhal, 1810) inordinata (J.Sahlberg, 1898) silesiaca (Kraatz, 1852) hybrida (Erichson, 1845) rugosa Stephens, 1829 brandisi Holdhaus, 1902 ciliaris (Schmidt, 1841) pallens (Sturm, 1807) triepkii (Schmidt, 1841) rubiginosa (Schmidt, 1841) polita (Marsham, 1802) calcarata (Erichson, 1845) lucens (Fairmaire, 1855) cinnamomea (Panzer, 1793) oblonga (Erichson, 1845) fracta (Seidlitz, 1874) rhaetica auct. nec (Erichson, 1845) FKSND-AI F-SN---- F-S-DEA- F------- FKSND--I -K------ FKSND-A- F-S-D--- F-SN----??--D--- FKSN--A-